diff --git a/webapp/themes/vitro/templates/footer.ftl b/webapp/themes/vitro/templates/footer.ftl
index fc258004a..29e054ead 100644
--- a/webapp/themes/vitro/templates/footer.ftl
+++ b/webapp/themes/vitro/templates/footer.ftl
@@ -7,25 +7,25 @@
         <#if copyright??>
             <small>&copy;${copyright.year?c}
             <#if copyright.url??>
-                <a href="${copyright.url}">${copyright.text}</a>
+                <a href="${copyright.url}" title="copyright">${copyright.text}</a>
             <#else>
                 ${copyright.text}
             </#if>
-             | <a class="terms" href="${urls.termsOfUse}">Terms of Use</a></small> | 
+             | <a class="terms" href="${urls.termsOfUse}" title="terms of use">Terms of Use</a></small> | 
         </#if>
         Powered by <a class="powered-by-vitro" href="http://vitro.sourceforge.net"><strong>Vitro</strong></a>
         <#if user.hasRevisionInfoAccess>
-             | Version <a href="${version.moreInfoUrl}">${version.label}</a>
+             | Version <a href="${version.moreInfoUrl}" title="version">${version.label}</a>
         </#if>
     </p>
     
     <nav role="navigation">
         <ul id="footer-nav" role="list">
-            <li role="listitem"><a href="${urls.about}">About</a></li>
+            <li role="listitem"><a href="${urls.about}" title="about">About</a></li>
             <#if urls.contact??>
-                <li role="listitem"><a href="${urls.contact}">Contact Us</a></li>
+                <li role="listitem"><a href="${urls.contact}" title="contact us">Contact Us</a></li>
             </#if> 
-            <li role="listitem"><a href="http://www.vivoweb.org/support" target="blank">Support</a></li>
+            <li role="listitem"><a href="http://www.vivoweb.org/support" target="blank" title="support">Support</a></li>
         </ul>
     </nav>
 </footer>
diff --git a/webapp/themes/vitro/templates/identity.ftl b/webapp/themes/vitro/templates/identity.ftl
index 59c666041..f102bd740 100644
--- a/webapp/themes/vitro/templates/identity.ftl
+++ b/webapp/themes/vitro/templates/identity.ftl
@@ -9,19 +9,19 @@
 
     <nav role="navigation">
         <ul id="header-nav" role="list">
-            <li role="listitem"><a href="${urls.index}">Index</a></li>
+            <li role="listitem"><a href="${urls.index}" title="index">Index</a></li>
             <#if user.loggedIn>
                 <#if user.hasSiteAdminAccess>
-                    <li role="listitem"><a href="${urls.siteAdmin}">Site Admin</a></li>
+                    <li role="listitem"><a href="${urls.siteAdmin}" title="site admin">Site Admin</a></li>
                 </#if>
                     <li>
                         <ul class="dropdown">
-                            <li id="user-menu"><a href="#">${user.loginName}</a>
+                            <li id="user-menu"><a href="#" title="user">${user.loginName}</a>
                                 <ul class="sub_menu">
                                      <#if urls.myAccount??>
-                                         <li role="listitem"><a href="${urls.myAccount}">My account</a></li>
+                                         <li role="listitem"><a href="${urls.myAccount}" title="my account">My account</a></li>
                                      </#if>
-                                     <li role="listitem"><a href="${urls.logout}">Log out</a></li>
+                                     <li role="listitem"><a href="${urls.logout}" title="log out">Log out</a></li>
                                 </ul>
                             </li>
                          </ul>
diff --git a/webapp/themes/vitro/templates/menu.ftl b/webapp/themes/vitro/templates/menu.ftl
index 03cd15766..a09f090d5 100644
--- a/webapp/themes/vitro/templates/menu.ftl
+++ b/webapp/themes/vitro/templates/menu.ftl
@@ -23,6 +23,6 @@
     
     <!--[if lte IE 8]>
     <noscript>
-        <p class="ie-alert">This site uses HTML elements that are not recognized by Internet Explorer 8 and below in the absence of JavaScript. As a result, the site will not be rendered appropriately. To correct this, please either enable JavaScript, upgrade to Internet Explorer 9, or use another browser. Here are the <a href="http://www.enable-javascript.com">instructions for enabling JavaScript in your web browser</a>.</p>
+        <p class="ie-alert">This site uses HTML elements that are not recognized by Internet Explorer 8 and below in the absence of JavaScript. As a result, the site will not be rendered appropriately. To correct this, please either enable JavaScript, upgrade to Internet Explorer 9, or use another browser. Here are the <a href="http://www.enable-javascript.com" title="javascript instructions">instructions for enabling JavaScript in your web browser</a>.</p>
     </noscript>
     <![endif]-->
\ No newline at end of file
diff --git a/webapp/web/templates/freemarker/body/accounts/userAccounts-acctCreatedEmail.ftl b/webapp/web/templates/freemarker/body/accounts/userAccounts-acctCreatedEmail.ftl
index b33469095..ec2bb70e4 100644
--- a/webapp/web/templates/freemarker/body/accounts/userAccounts-acctCreatedEmail.ftl
+++ b/webapp/web/templates/freemarker/body/accounts/userAccounts-acctCreatedEmail.ftl
@@ -32,7 +32,7 @@
         </p>
         
         <p>
-            <a href="${passwordLink}">${passwordLink}</a>
+            <a href="${passwordLink}" title="password">${passwordLink}</a>
         </p>
         
         <p>
diff --git a/webapp/web/templates/freemarker/body/accounts/userAccounts-add.ftl b/webapp/web/templates/freemarker/body/accounts/userAccounts-add.ftl
index b3e1ffec5..bb0a3c5f1 100644
--- a/webapp/web/templates/freemarker/body/accounts/userAccounts-add.ftl
+++ b/webapp/web/templates/freemarker/body/accounts/userAccounts-add.ftl
@@ -2,7 +2,7 @@
 
 <#-- Template for adding a user account -->
 
-<h3><a class="account-menu" href="accountsAdmin">User accounts</a> > Add new account</h3>
+<h3><a class="account-menu" href="accountsAdmin" title="add new account">User accounts</a> > Add new account</h3>
 
     <#if errorEmailIsEmpty??>
         <#assign errorMessage = "You must supply an email address." />
@@ -89,7 +89,7 @@
             </section>
         </#if>
         
-        <p><input type="submit" name="submitAdd" value="Add new account" class="submit" /> or <a class="cancel" href="${formUrls.list}">Cancel</a></p>
+        <p><input type="submit" name="submitAdd" value="Add new account" class="submit" /> or <a class="cancel" href="${formUrls.list}" title="cancel">Cancel</a></p>
 
         <p class="requiredHint">* required fields</p>
     </form>
diff --git a/webapp/web/templates/freemarker/body/accounts/userAccounts-associateProfilePanel.ftl b/webapp/web/templates/freemarker/body/accounts/userAccounts-associateProfilePanel.ftl
index 0f8beba2c..f75ef2856 100644
--- a/webapp/web/templates/freemarker/body/accounts/userAccounts-associateProfilePanel.ftl
+++ b/webapp/web/templates/freemarker/body/accounts/userAccounts-associateProfilePanel.ftl
@@ -26,8 +26,8 @@ ${stylesheets.add('<link rel="stylesheet" href="${urls.base}/edit/forms/css/auto
             <p>
                 <label for="associatedProfileName">Associated profile:</label>
                 <span class="acSelectionInfo" id="associatedProfileName"></span>
-                <a href="" id="verifyProfileLink">(verify this match)</a>
-                <a href="" id="changeProfileLink">(change profile)</a>
+                <a href="" id="verifyProfileLink" title="verify this match">(verify this match)</a>
+                <a href="" id="changeProfileLink" title="change profile">(change profile)</a>
             </p>
             <input type="hidden" id="associatedProfileUri" name="associatedProfileUri" value="" />
         </div>
diff --git a/webapp/web/templates/freemarker/body/accounts/userAccounts-edit.ftl b/webapp/web/templates/freemarker/body/accounts/userAccounts-edit.ftl
index 99c95c124..d1a826042 100644
--- a/webapp/web/templates/freemarker/body/accounts/userAccounts-edit.ftl
+++ b/webapp/web/templates/freemarker/body/accounts/userAccounts-edit.ftl
@@ -2,7 +2,7 @@
 
 <#-- Template for editing a user account -->
 
-<h3><a class="account-menu" href="accountsAdmin">User accounts</a> > Edit account</h3>
+<h3><a class="account-menu" href="accountsAdmin" title="edit account">User accounts</a> > Edit account</h3>
 
     <#if errorEmailIsEmpty??>
         <#assign errorMessage = "You must supply an email address." />
@@ -97,7 +97,7 @@
             </section>
         </#if>
 
-        <p><input type="submit" id="submitMyAccount" name="submitEdit" value="Save changes" class="submit" disabled /> or <a class="cancel" href="${formUrls.list}">Cancel</a></p>
+        <p><input type="submit" id="submitMyAccount" name="submitEdit" value="Save changes" class="submit" disabled /> or <a class="cancel" href="${formUrls.list}" title="cancel">Cancel</a></p>
 
         <p class="requiredHint">* required fields</p>
     </form>
diff --git a/webapp/web/templates/freemarker/body/accounts/userAccounts-firstTimeExternal.ftl b/webapp/web/templates/freemarker/body/accounts/userAccounts-firstTimeExternal.ftl
index f8885fa3c..a1f725bca 100644
--- a/webapp/web/templates/freemarker/body/accounts/userAccounts-firstTimeExternal.ftl
+++ b/webapp/web/templates/freemarker/body/accounts/userAccounts-firstTimeExternal.ftl
@@ -54,7 +54,7 @@
             </p>
         </#if>
 
-        <p><input type="submit" name="submit" value="Create account" class="submit"/> or <a class="cancel" href="${urls.home}">Cancel</a></p>
+        <p><input type="submit" name="submit" value="Create account" class="submit"/> or <a class="cancel" href="${urls.home}" title="cancel">Cancel</a></p>
     </form>
 </section>
 
diff --git a/webapp/web/templates/freemarker/body/accounts/userAccounts-list.ftl b/webapp/web/templates/freemarker/body/accounts/userAccounts-list.ftl
index 8ea73f014..2194c0650 100644
--- a/webapp/web/templates/freemarker/body/accounts/userAccounts-list.ftl
+++ b/webapp/web/templates/freemarker/body/accounts/userAccounts-list.ftl
@@ -12,7 +12,7 @@
     <section class="account-feedback">
         <p>
             A new account for 
-            <a href="${newUserAccount.editUrl}">${newUserAccount.firstName} ${newUserAccount.lastName}</a>
+            <a href="${newUserAccount.editUrl}" title="new account">${newUserAccount.firstName} ${newUserAccount.lastName}</a>
             was successfully created. 
             <#if emailWasSent?? > 
                 A notification email has been sent to ${newUserAccount.emailAddress}
@@ -26,7 +26,7 @@
     <section class="account-feedback">
         <p>
             The account for 
-            <a href="${updatedUserAccount.editUrl}">${updatedUserAccount.firstName} ${updatedUserAccount.lastName}</a>
+            <a href="${updatedUserAccount.editUrl}" title="updated account">${updatedUserAccount.firstName} ${updatedUserAccount.lastName}</a>
             has been updated.
             <#if emailWasSent?? > 
                 A confirmation email has been sent to ${updatedUserAccount.emailAddress}
@@ -60,7 +60,7 @@
         </select>
         
         <#if roleFilterUri?has_content>
-             <span><a href="${formUrls.list}"> View all accounts</a></span>
+             <span><a href="${formUrls.list}" title="view all acounts"> View all accounts</a></span>
         </#if>
     </form>
 </section>
@@ -80,7 +80,7 @@
 </section>
 <#if searchTerm?has_content>
 <section id="search-feedback">
-    <p>Search results for "<strong>${searchTerm}</strong>" | <span><a href="${formUrls.list}"> View all accounts</a></span></p>
+    <p>Search results for "<strong>${searchTerm}</strong>" | <span><a href="${formUrls.list}" title="view all accounts"> View all accounts</a></span></p>
 </section>
 </#if>
 <form method="POST" action="${formUrls.list}" id="account-display" class="customForm" role="accounts display">
@@ -156,7 +156,7 @@
                         </#if>
     
                         <#if account.editUrl != "">
-                            <a ${disableDeleteAccount} href="${account.editUrl}" >${account.emailAddress}</a>
+                            <a ${disableDeleteAccount} href="${account.editUrl}"  title="disable account">${account.emailAddress}</a>
                             <!-- when this link is clicked, editAccount is noticed and all other fields are ignored. -->
                         <#else>
                             ${account.emailAddress}
diff --git a/webapp/web/templates/freemarker/body/accounts/userAccounts-myAccount.ftl b/webapp/web/templates/freemarker/body/accounts/userAccounts-myAccount.ftl
index e77b1b52b..27add004a 100644
--- a/webapp/web/templates/freemarker/body/accounts/userAccounts-myAccount.ftl
+++ b/webapp/web/templates/freemarker/body/accounts/userAccounts-myAccount.ftl
@@ -84,7 +84,7 @@
             <input type="password" name="confirmPassword" value="${confirmPassword}" id="confirm-password" role="input" />
         </#if>
 
-        <p><input type="submit" id="submitMyAccount" name="submitMyAccount" value="Save changes" class="submit" disabled /> or <a class="cancel" href="${urls.referringPage}">Cancel</a></p>
+        <p><input type="submit" id="submitMyAccount" name="submitMyAccount" value="Save changes" class="submit" disabled /> or <a class="cancel" href="${urls.referringPage}" title="cancel">Cancel</a></p>
 
         <p class="requiredHint">* required fields</p>
     </form>
diff --git a/webapp/web/templates/freemarker/body/accounts/userAccounts-myProxiesPanel.ftl b/webapp/web/templates/freemarker/body/accounts/userAccounts-myProxiesPanel.ftl
index 195d9d5de..5481ca409 100644
--- a/webapp/web/templates/freemarker/body/accounts/userAccounts-myProxiesPanel.ftl
+++ b/webapp/web/templates/freemarker/body/accounts/userAccounts-myProxiesPanel.ftl
@@ -35,7 +35,7 @@
                 
                 <p class="proxy-info">%label% | <span class="class-label">%classLabel%</span>
                     <br />
-                    <a class='remove-proxy' href="." templatePart="remove">Remove selection</a>
+                    <a class='remove-proxy' href="." templatePart="remove" title="remove selection">Remove selection</a>
                     
                     <input type="hidden" name="proxyUri" value="%uri%" role="input" />
                 </p>
diff --git a/webapp/web/templates/freemarker/body/admin/startupStatus-displayRaw.ftl b/webapp/web/templates/freemarker/body/admin/startupStatus-displayRaw.ftl
index e9bb2194e..a8db53dc1 100644
--- a/webapp/web/templates/freemarker/body/admin/startupStatus-displayRaw.ftl
+++ b/webapp/web/templates/freemarker/body/admin/startupStatus-displayRaw.ftl
@@ -100,7 +100,7 @@
             
             <#-- If there were no fatal errors, let them go forward from here. -->
             <#if showLink>
-                <p><a href=".">Continue</a></p>
+                <p><a href="." title="continue">Continue</a></p>
     	    </#if>
             
         </#if>
diff --git a/webapp/web/templates/freemarker/body/classGroups.ftl b/webapp/web/templates/freemarker/body/classGroups.ftl
index aff2f032d..9d12507d7 100644
--- a/webapp/web/templates/freemarker/body/classGroups.ftl
+++ b/webapp/web/templates/freemarker/body/classGroups.ftl
@@ -16,7 +16,7 @@
                             <#list classGroup.classes as class> 
                                 <#-- Only render populated classes -->
                                 <#if (class.individualCount > 0)>
-                                    <li role="listitem"><a href="${class.url}">${class.name}</a> (${class.individualCount})</li>
+                                    <li role="listitem"><a href="${class.url}" title="class name">${class.name}</a> (${class.individualCount})</li>
                                 </#if>
                             </#list>
                         </ul>
diff --git a/webapp/web/templates/freemarker/body/contactForm/contactForm-error.ftl b/webapp/web/templates/freemarker/body/contactForm/contactForm-error.ftl
index 7027d14e3..93bfbd4cb 100644
--- a/webapp/web/templates/freemarker/body/contactForm/contactForm-error.ftl
+++ b/webapp/web/templates/freemarker/body/contactForm/contactForm-error.ftl
@@ -10,4 +10,4 @@
     </section>
 </#if>
 
-<p class="contactUsReturnHome">Return to the <a href="${urls.home}">home page</a>.</p> 
\ No newline at end of file
+<p class="contactUsReturnHome">Return to the <a href="${urls.home}" title="home page">home page</a>.</p> 
\ No newline at end of file
diff --git a/webapp/web/templates/freemarker/body/error/error-display.ftl b/webapp/web/templates/freemarker/body/error/error-display.ftl
index ebfc6ab7e..bd1936097 100644
--- a/webapp/web/templates/freemarker/body/error/error-display.ftl
+++ b/webapp/web/templates/freemarker/body/error/error-display.ftl
@@ -24,6 +24,6 @@
     </#if>  
 
 <#elseif ! errorOnHomePage> <#-- view for other users -->
-    <p>Return to the <a href="${urls.home}">home page</a></p> 
+    <p>Return to the <a href="${urls.home}" title="home page">home page</a></p> 
 </#if>
 
diff --git a/webapp/web/templates/freemarker/body/error/error-standard.ftl b/webapp/web/templates/freemarker/body/error/error-standard.ftl
index 1403c8997..1eedaf780 100644
--- a/webapp/web/templates/freemarker/body/error/error-standard.ftl
+++ b/webapp/web/templates/freemarker/body/error/error-standard.ftl
@@ -4,4 +4,4 @@
 
 <p>There was an error in the system.</p>
 
-<p>Return to the <a href="${urls.home}">home page</a>.</p> 
\ No newline at end of file
+<p>Return to the <a href="${urls.home}" title="home page">home page</a>.</p> 
\ No newline at end of file
diff --git a/webapp/web/templates/freemarker/body/imageUpload/imageUpload-cropImage.ftl b/webapp/web/templates/freemarker/body/imageUpload/imageUpload-cropImage.ftl
index 7dc768ce7..9f6f94ad0 100644
--- a/webapp/web/templates/freemarker/body/imageUpload/imageUpload-cropImage.ftl
+++ b/webapp/web/templates/freemarker/body/imageUpload/imageUpload-cropImage.ftl
@@ -45,7 +45,7 @@ ${stylesheets.add('<link rel="stylesheet" href="${urls.base}/css/uploadImages.cs
                                       
                 <input  class="submit" type="submit" value="Save photo">
                 
-                <span class="or"> or <a class="cancel"  href="${cancelUrl}">Cancel</a></span>
+                <span class="or"> or <a class="cancel"  href="${cancelUrl}" title="cancel">Cancel</a></span>
             </form>
        </section>
     </section>
diff --git a/webapp/web/templates/freemarker/body/imageUpload/imageUpload-newImage.ftl b/webapp/web/templates/freemarker/body/imageUpload/imageUpload-newImage.ftl
index 2c1d6ffa8..675b63920 100644
--- a/webapp/web/templates/freemarker/body/imageUpload/imageUpload-newImage.ftl
+++ b/webapp/web/templates/freemarker/body/imageUpload/imageUpload-newImage.ftl
@@ -30,6 +30,6 @@ ${stylesheets.add('<link rel="stylesheet" href="${urls.base}/css/uploadImages.cs
         Minimum image dimensions: ${thumbnailWidth} x ${thumbnailHeight} pixels</p>
         <input class="submit" type="submit" value="Upload photo"/>
         
-        <span class="or"> or <a class="cancel"  href="${cancelUrl}">Cancel</a></span>
+        <span class="or"> or <a class="cancel"  href="${cancelUrl}" title="cancel">Cancel</a></span>
     </form>
 </section>
\ No newline at end of file
diff --git a/webapp/web/templates/freemarker/body/imageUpload/imageUpload-replaceImage.ftl b/webapp/web/templates/freemarker/body/imageUpload/imageUpload-replaceImage.ftl
index 06f501ebf..d8850d188 100644
--- a/webapp/web/templates/freemarker/body/imageUpload/imageUpload-replaceImage.ftl
+++ b/webapp/web/templates/freemarker/body/imageUpload/imageUpload-replaceImage.ftl
@@ -31,6 +31,6 @@ ${stylesheets.add('<link rel="stylesheet" href="${urls.base}/css/uploadImages.cs
         Minimum image dimensions: ${thumbnailWidth}x${thumbnailHeight} pixels</p>
         <input class="submit" type="submit" value="Upload photo"> 
 
-        <span class="or"> or <a class="cancel"  href="${cancelUrl}">Cancel</a></span>
+        <span class="or"> or <a class="cancel"  href="${cancelUrl}" title="cancel">Cancel</a></span>
      </form>
 </section>
\ No newline at end of file
diff --git a/webapp/web/templates/freemarker/body/individual/individual-vitro.ftl b/webapp/web/templates/freemarker/body/individual/individual-vitro.ftl
index fa2ae5655..61182a21b 100644
--- a/webapp/web/templates/freemarker/body/individual/individual-vitro.ftl
+++ b/webapp/web/templates/freemarker/body/individual/individual-vitro.ftl
@@ -25,7 +25,7 @@
         <header>
             <#if relatedSubject??>
                 <h2>${relatedSubject.relatingPredicateDomainPublic} for ${relatedSubject.name}</h2>
-                <p><a href="${relatedSubject.url}">&larr; return to ${relatedSubject.name}</a></p>                
+                <p><a href="${relatedSubject.url}" title="return to subject">&larr; return to ${relatedSubject.name}</a></p>                
             <#else>                
                 <h1 class="fn">
                     <#-- Label -->
diff --git a/webapp/web/templates/freemarker/body/individualList.ftl b/webapp/web/templates/freemarker/body/individualList.ftl
index bfa0300aa..41a9065bd 100644
--- a/webapp/web/templates/freemarker/body/individualList.ftl
+++ b/webapp/web/templates/freemarker/body/individualList.ftl
@@ -31,7 +31,7 @@ ${stylesheets.add('<link rel="stylesheet" href="${urls.base}/css/browseIndex.css
                                 <li class="selectedNavPage">${page.text}</li>
                             <#else>
                                 <#-- RY Ideally the urls would be generated by the controller; see search-pagedResults.ftl -->
-                                <li><a href="${urls.base}/individuallist?${page.param}&vclassId=${vclassId?url}">${page.text}</a></li>
+                                <li><a href="${urls.base}/individuallist?${page.param}&vclassId=${vclassId?url}" title="page text">${page.text}</a></li>
                             </#if>
                         </#list>
                     </ul>
diff --git a/webapp/web/templates/freemarker/body/manageproxies/manageProxies-list.ftl b/webapp/web/templates/freemarker/body/manageproxies/manageProxies-list.ftl
index 1aa954d14..16a241e86 100644
--- a/webapp/web/templates/freemarker/body/manageproxies/manageProxies-list.ftl
+++ b/webapp/web/templates/freemarker/body/manageproxies/manageProxies-list.ftl
@@ -48,7 +48,7 @@
                             <p class="proxy-info">
                                 %label% | <span class="class-label">%classLabel%</span>
                                 <br />
-                                <a class='remove-proxy' href="." templatePart="remove">Remove selection</a>
+                                <a class='remove-proxy' href="." templatePart="remove" title="remove selection">Remove selection</a>
                                 <input type="hidden" name="proxyUri" value="%uri%" >
                             </p>
                         </li>
@@ -79,7 +79,7 @@
                             <img class="photo-profile" width="60" alt="%label%" src="%imageUrl%">
                             
                             <p class="proxy-info-profile">%label% | <span class="class-label">%classLabel%</span>
-                                <br /><a class='remove-proxy' href="." templatePart="remove">Remove selection</a>
+                                <br /><a class='remove-proxy' href="." templatePart="remove" title="remove selection">Remove selection</a>
                             </p>
     
                             <input type="hidden" name="profileUri" templatePart="uriField" value="%uri%" role="input" />
@@ -101,7 +101,7 @@
         <input type="text" name="searchTerm" role="input" />
         <input class="submit" type="submit" name="searchByProxy" value="Search" role="button" /> 
             <#if page.previous??>
-               | <a href="${formUrls.list}?pageIndex=${page.previous}&searchTerm=${searchTerm}">Previous</a>
+               | <a href="${formUrls.list}?pageIndex=${page.previous}&searchTerm=${searchTerm}" title="previous">Previous</a>
             </#if>
             
             <#if page.last != 0>
@@ -109,11 +109,11 @@
             </#if>
             
             <#if page.next??>
-                <a href="${formUrls.list}?pageIndex=${page.next}&searchTerm=${searchTerm}">Next</a>
+                <a href="${formUrls.list}?pageIndex=${page.next}&searchTerm=${searchTerm}" title="next">Next</a>
             </#if>
 
             <#if searchTerm?has_content>
-                <p>Search results for "<span class="blue">${searchTerm}</span>" | <a href="${formUrls.list}">View all profile editors</a></p>
+                <p>Search results for "<span class="blue">${searchTerm}</span>" | <a href="${formUrls.list}" title="view all profile editors">View all profile editors</a></p>
             </#if>
     </form>
 </section>
@@ -129,7 +129,7 @@
                 <p class="proxyInfoElement proxy-info">
                     ${p.label} | <span class="class-label">${p.classLabel}</span>
                     <br>
-                    <a class="remove-proxyUri" href="${formUrls.edit}?proxyUri=${p.uri}&deleteProxy=Delete proxy">Delete profile editor</a>
+                    <a class="remove-proxyUri" href="${formUrls.edit}?proxyUri=${p.uri}&deleteProxy=Delete proxy" title="delete profile editor">Delete profile editor</a>
                     <input type="hidden" value="${p.uri}" name="proxyUri">
                 </p>
             </div>       
@@ -168,7 +168,7 @@
                             <img class="photo-profile" width="60" alt="%label%" src="%imageUrl%">
                              
                             <p class="proxy-info-profile">%label% | <span class="class-label">%classLabel%</span>
-                                <br /><a class='remove-proxy' href="." templatePart="remove">Remove selection</a>
+                                <br /><a class='remove-proxy' href="." templatePart="remove" title="remove selection">Remove selection</a>
                             </p>
                         </li>
                         
diff --git a/webapp/web/templates/freemarker/body/menuN3Edit.ftl b/webapp/web/templates/freemarker/body/menuN3Edit.ftl
index eb6ad2c52..f9f3ea212 100644
--- a/webapp/web/templates/freemarker/body/menuN3Edit.ftl
+++ b/webapp/web/templates/freemarker/body/menuN3Edit.ftl
@@ -20,7 +20,7 @@
             <textarea name="navigationN3" id="navigationN3" cols="45" rows="40" class="maxWidth">
                 ${menuN3}<#t><#-- The trim directive here is to trim leading and trailing white-space -->
             </textarea><#lt><#-- This directive trims only leading white-space -->
-            <input name="submit" id="submit" value="Save" type="submit"/> or <a href="${urls.base}${cancelUrl}">Cancel</a>
+            <input name="submit" id="submit" value="Save" type="submit"/> or <a href="${urls.base}${cancelUrl}" title="cancel">Cancel</a>
         </form>
     </#if>
 </div>
\ No newline at end of file
diff --git a/webapp/web/templates/freemarker/body/partials/account/userAccounts-accountsNav.ftl b/webapp/web/templates/freemarker/body/partials/account/userAccounts-accountsNav.ftl
index d9efc2f62..2b2f75f1d 100644
--- a/webapp/web/templates/freemarker/body/partials/account/userAccounts-accountsNav.ftl
+++ b/webapp/web/templates/freemarker/body/partials/account/userAccounts-accountsNav.ftl
@@ -35,11 +35,11 @@
           accounts per page <input type="submit" name="accounts-per-page" value="Update" /> | 
 
           <#if page.previous?has_content>
-              <a href="${formUrls.list}?accountsPerPage=${accountsPerPage}&pageIndex=${page.previous}">Previous</a> <!-- only present if current page is not 1.-->
+              <a href="${formUrls.list}?accountsPerPage=${accountsPerPage}&pageIndex=${page.previous}" title="previous">Previous</a> <!-- only present if current page is not 1.-->
           </#if>
               ${page.current} of ${page.last} 
           <#if page.next?has_content>
-              <a href="${formUrls.list}?accountsPerPage=${accountsPerPage}&pageIndex=${page.next}">Next</a><!-- only present if current page is not last page.-->
+              <a href="${formUrls.list}?accountsPerPage=${accountsPerPage}&pageIndex=${page.next}" title="next">Next</a><!-- only present if current page is not last page.-->
           </#if>
       </nav>
   </section>
diff --git a/webapp/web/templates/freemarker/body/partials/class/view/search/view-search-default.ftl b/webapp/web/templates/freemarker/body/partials/class/view/search/view-search-default.ftl
index e74adaab9..05273515e 100644
--- a/webapp/web/templates/freemarker/body/partials/class/view/search/view-search-default.ftl
+++ b/webapp/web/templates/freemarker/body/partials/class/view/search/view-search-default.ftl
@@ -4,7 +4,7 @@
 
 <#import "lib-properties.ftl" as p>
 
-<a href="${individual.profileUrl}">${individual.name}</a>
+<a href="${individual.profileUrl}" title="name">${individual.name}</a>
 
 <@p.mostSpecificTypes individual />
 
diff --git a/webapp/web/templates/freemarker/body/partials/individual/individual-adminPanel.ftl b/webapp/web/templates/freemarker/body/partials/individual/individual-adminPanel.ftl
index f68ad9ba6..175269b54 100644
--- a/webapp/web/templates/freemarker/body/partials/individual/individual-adminPanel.ftl
+++ b/webapp/web/templates/freemarker/body/partials/individual/individual-adminPanel.ftl
@@ -6,7 +6,7 @@
 
 <#if individual.showAdminPanel>
     <section id="admin">
-        <h3>Admin Panel</h3><a class="edit-individual" href="${individual.controlPanelUrl()}">Edit this individual</a>
+        <h3>Admin Panel</h3><a class="edit-individual" href="${individual.controlPanelUrl()}" title="edit this individual">Edit this individual</a>
         
         <section id = "verbose-mode">
         <#if verbosePropertySwitch?has_content>
@@ -14,10 +14,10 @@
             <#assign currentValue = verbosePropertySwitch.currentValue?string("on", "off")>
             <#assign newValue = verbosePropertySwitch.currentValue?string("off", "on")>
             <span>Verbose property display is <b>${currentValue}</b> | </span>
-            <a id="${anchorId}" class="verbose-toggle small" href="${verbosePropertySwitch.url}#${anchorId}">Turn ${newValue}</a>
+            <a id="${anchorId}" class="verbose-toggle small" href="${verbosePropertySwitch.url}#${anchorId}" title="verbose control">Turn ${newValue}</a>
         </#if> 
         </section>
 
-        <p class="uri-link">Resource URI: <a href="${individual.uri}" target="_blank">${individual.uri}</a></p>
+        <p class="uri-link">Resource URI: <a href="${individual.uri}" target="_blank" title="resource uri">${individual.uri}</a></p>
     </section>
 </#if>
\ No newline at end of file
diff --git a/webapp/web/templates/freemarker/body/partials/individual/individual-properties.ftl b/webapp/web/templates/freemarker/body/partials/individual/individual-properties.ftl
index b7c3be07e..fc322ab25 100644
--- a/webapp/web/templates/freemarker/body/partials/individual/individual-properties.ftl
+++ b/webapp/web/templates/freemarker/body/partials/individual/individual-properties.ftl
@@ -9,7 +9,7 @@
     
     <section class="property-group" role="region">
         <nav class="scroll-up" role="navigation">
-            <a href="#branding">
+            <a href="#branding" title="scroll up">
                 <img src="${urls.images}/individual/scroll-up.gif" alt="scroll to property group menus" />
             </a>
         </nav>
diff --git a/webapp/web/templates/freemarker/body/partials/individual/individual-propertyGroupMenu.ftl b/webapp/web/templates/freemarker/body/partials/individual/individual-propertyGroupMenu.ftl
index 58cc4a81a..04fec93b7 100644
--- a/webapp/web/templates/freemarker/body/partials/individual/individual-propertyGroupMenu.ftl
+++ b/webapp/web/templates/freemarker/body/partials/individual/individual-propertyGroupMenu.ftl
@@ -16,7 +16,7 @@
                         <#if groupname?has_content>
                             <#-- capitalize will capitalize each word in the name; cap_first only the first. We may need a custom
                             function to capitalize all except function words. -->
-                            <li role="listitem"><a href="#${groupname}">${groupname?capitalize}</a></li>
+                            <li role="listitem"><a href="#${groupname}" title="group name">${groupname?capitalize}</a></li>
                         </#if>
                     </#list>
                 </ul>
diff --git a/webapp/web/templates/freemarker/body/partials/individual/propStatement-default.ftl b/webapp/web/templates/freemarker/body/partials/individual/propStatement-default.ftl
index dee213299..b06146531 100644
--- a/webapp/web/templates/freemarker/body/partials/individual/propStatement-default.ftl
+++ b/webapp/web/templates/freemarker/body/partials/individual/propStatement-default.ftl
@@ -6,4 +6,4 @@
      is also used to generate the property statement during a deletion.  
  -->
 
-<a href="${profileUrl(statement.object)}">${statement.label!statement.localName!}</a> 
+<a href="${profileUrl(statement.object)}" title="name">${statement.label!statement.localName!}</a> 
diff --git a/webapp/web/templates/freemarker/body/partials/individual/propStatement-simple.ftl b/webapp/web/templates/freemarker/body/partials/individual/propStatement-simple.ftl
index 1c870b613..1419ce359 100644
--- a/webapp/web/templates/freemarker/body/partials/individual/propStatement-simple.ftl
+++ b/webapp/web/templates/freemarker/body/partials/individual/propStatement-simple.ftl
@@ -2,4 +2,4 @@
 
 <#-- Simple object property statement template -->
 
-<a href="${profileUrl(statement.object)}">${statement.name!}</a>  
+<a href="${profileUrl(statement.object)}" title="name">${statement.name!}</a>  
diff --git a/webapp/web/templates/freemarker/body/partials/listedIndividual.ftl b/webapp/web/templates/freemarker/body/partials/listedIndividual.ftl
index 725bed092..e732f00ab 100644
--- a/webapp/web/templates/freemarker/body/partials/listedIndividual.ftl
+++ b/webapp/web/templates/freemarker/body/partials/listedIndividual.ftl
@@ -4,6 +4,6 @@
 
 <#import "lib-properties.ftl" as p>
 
-<a href="${individual.profileUrl}">${individual.name}</a>
+<a href="${individual.profileUrl}" title="name">${individual.name}</a>
 
 <@p.mostSpecificTypes individual  />
\ No newline at end of file
diff --git a/webapp/web/templates/freemarker/body/partials/menupage/menupage-browse.ftl b/webapp/web/templates/freemarker/body/partials/menupage/menupage-browse.ftl
index c90253b1b..9a8c254aa 100644
--- a/webapp/web/templates/freemarker/body/partials/menupage/menupage-browse.ftl
+++ b/webapp/web/templates/freemarker/body/partials/menupage/menupage-browse.ftl
@@ -24,7 +24,7 @@
             <h3 class="selected-class"></h3>
             <#assign alphabet = ["A", "B", "C", "D", "E", "F", "G" "H", "I", "J", "K", "L", "M", "N", "O", "P", "Q", "R", "S", "T", "U", "V", "W", "X", "Y", "Z"] />
             <ul id="alpha-browse-individuals">
-                <li><a href="#" class="selected" data-alpha="all">All</a></li>
+                <li><a href="#" class="selected" data-alpha="all" title="select all">All</a></li>
                 <#list alphabet as letter>
                     <li><a href="#" data-alpha="${letter?lower_case}" title="Browse all individuals whose name starts with ${letter}">${letter}</a></li>
                 </#list>
diff --git a/webapp/web/templates/freemarker/body/search/search-pagedResults.ftl b/webapp/web/templates/freemarker/body/search/search-pagedResults.ftl
index f58fb4d89..70a094339 100644
--- a/webapp/web/templates/freemarker/body/search/search-pagedResults.ftl
+++ b/webapp/web/templates/freemarker/body/search/search-pagedResults.ftl
@@ -16,7 +16,7 @@
             <h4>Display only</h4>           
             <ul>           
             <#list classGroupLinks as link>
-                <li><a href="${link.url}">${link.text}</a></li>
+                <li><a href="${link.url}" title="class group link">${link.text}</a></li>
             </#list>
             </ul>           
         </div>
@@ -27,7 +27,7 @@
             <h4>Limit ${classGroupName} to</h4>
             <ul>           
             <#list classLinks as link>
-                <li><a href="${link.url}">${link.text}</a></li>
+                <li><a href="${link.url}" title="class link">${link.text}</a></li>
             </#list>
             </ul>
         </div>
@@ -46,15 +46,15 @@
     <#if (pagingLinks?size > 0)>
         <div class="searchpages">
             Pages: 
-            <#if prevPage??><a class="prev" href="${prevPage}">Previous</a></#if>
+            <#if prevPage??><a class="prev" href="${prevPage}" title="previous">Previous</a></#if>
             <#list pagingLinks as link>
                 <#if link.url??>
-                    <a href="${link.url}">${link.text}</a>
+                    <a href="${link.url}" title="page link">${link.text}</a>
                 <#else>
                     <span>${link.text}</span> <#-- no link if current page -->
                 </#if>
             </#list>
-            <#if nextPage??><a class="next" href="${nextPage}">Next</a></#if>
+            <#if nextPage??><a class="next" href="${nextPage}" title="next">Next</a></#if>
         </div>
     </#if>
 
diff --git a/webapp/web/templates/freemarker/body/siteAdmin/siteAdmin-advancedDataTools.ftl b/webapp/web/templates/freemarker/body/siteAdmin/siteAdmin-advancedDataTools.ftl
index 8aea6e511..807ce50fd 100644
--- a/webapp/web/templates/freemarker/body/siteAdmin/siteAdmin-advancedDataTools.ftl
+++ b/webapp/web/templates/freemarker/body/siteAdmin/siteAdmin-advancedDataTools.ftl
@@ -7,11 +7,11 @@
         <h3>Advanced Data Tools</h3>
         
         <ul role="navigation">
-            <li role="listitem"><a href="${dataTools.rdfData}">Add/Remove RDF data</a></li>
-            <li role="listitem"><a href="${dataTools.ingest}">Ingest tools</a></li>
-            <li role="listitem"><a href="${dataTools.rdfExport}">RDF export</a></li>
-            <li role="listitem"><a href="${dataTools.sparqlQuery}">SPARQL query</a></li>
-            <li role="listitem"><a href="${dataTools.sparqlQueryBuilder}">SPARQL query builder</a></li>
+            <li role="listitem"><a href="${dataTools.rdfData}" title="Add/Remove RDF data">Add/Remove RDF data</a></li>
+            <li role="listitem"><a href="${dataTools.ingest}" title="Ingest tools">Ingest tools</a></li>
+            <li role="listitem"><a href="${dataTools.rdfExport}" title="RDF export">RDF export</a></li>
+            <li role="listitem"><a href="${dataTools.sparqlQuery}" title="SPARQL query">SPARQL query</a></li>
+            <li role="listitem"><a href="${dataTools.sparqlQueryBuilder}" title="SPARQL query builder">SPARQL query builder</a></li>
         </ul>
     </section>
 </#if>
\ No newline at end of file
diff --git a/webapp/web/templates/freemarker/body/siteAdmin/siteAdmin-indexCacheRebuild.ftl b/webapp/web/templates/freemarker/body/siteAdmin/siteAdmin-indexCacheRebuild.ftl
index 8bd127812..97d72da42 100644
--- a/webapp/web/templates/freemarker/body/siteAdmin/siteAdmin-indexCacheRebuild.ftl
+++ b/webapp/web/templates/freemarker/body/siteAdmin/siteAdmin-indexCacheRebuild.ftl
@@ -8,19 +8,19 @@
         
         <ul role="navigation">
             <#if indexCacheRebuild.rebuildClassGroupCache?has_content>
-                <li role="listitem"><a href="${indexCacheRebuild.rebuildClassGroupCache}">Rebuild class group cache</a></li>
+                <li role="listitem"><a href="${indexCacheRebuild.rebuildClassGroupCache}" title="Rebuild class group cache">Rebuild class group cache</a></li>
             </#if>
             
             <#if indexCacheRebuild.rebuildSearchIndex?has_content>
-                <li role="listitem"><a href="${indexCacheRebuild.rebuildSearchIndex }">Rebuild search index</a></li>
+                <li role="listitem"><a href="${indexCacheRebuild.rebuildSearchIndex }" title="Rebuild search index">Rebuild search index</a></li>
             </#if>
             
             <#if indexCacheRebuild.rebuildVisCache?has_content>
-                <li role="listitem"><a href="${indexCacheRebuild.rebuildVisCache}">Rebuild visualization cache</a></li>
+                <li role="listitem"><a href="${indexCacheRebuild.rebuildVisCache}" title="Rebuild visualization cache">Rebuild visualization cache</a></li>
             </#if>
             
             <#if indexCacheRebuild.recomputeInferences?has_content>
-                <li role="listitem"><a href="${indexCacheRebuild.recomputeInferences}">Recompute inferences</a></li>
+                <li role="listitem"><a href="${indexCacheRebuild.recomputeInferences}" title="Recompute inferences">Recompute inferences</a></li>
             </#if>
         </ul>
     </section>
diff --git a/webapp/web/templates/freemarker/body/siteAdmin/siteAdmin-ontologyEditor.ftl b/webapp/web/templates/freemarker/body/siteAdmin/siteAdmin-ontologyEditor.ftl
index e6e891992..db61bdd30 100644
--- a/webapp/web/templates/freemarker/body/siteAdmin/siteAdmin-ontologyEditor.ftl
+++ b/webapp/web/templates/freemarker/body/siteAdmin/siteAdmin-ontologyEditor.ftl
@@ -18,19 +18,19 @@
         <h4>Class Management</h4>
         
         <ul role="navigation">
-            <li role="listitem"><a href="${ontologyEditor.urls.classGroups}">Class groups</a></li>
-            <li role="listitem"><a href="${ontologyEditor.urls.classHierarchy}">Class hierarchy</a></li>
+            <li role="listitem"><a href="${ontologyEditor.urls.classGroups}" title="Class groups">Class groups</a></li>
+            <li role="listitem"><a href="${ontologyEditor.urls.classHierarchy}" title="Class hierarchy">Class hierarchy</a></li>
             
         </ul>
         
         <h4>Property Management</h4>
         
         <ul role="navigation">
-            <li role="listitem"><a href="${ontologyEditor.urls.dataPropertyHierarchy}">Data property hierarchy</a></li>
-            <li role="listitem"><a href="${ontologyEditor.urls.objectPropertyHierarchy}">Object property hierarchy</a></li>
-            <li role="listitem"><a href="${ontologyEditor.urls.propertyGroups}">Property groups</a></li>
+            <li role="listitem"><a href="${ontologyEditor.urls.dataPropertyHierarchy}" title="Data property hierarchy">Data property hierarchy</a></li>
+            <li role="listitem"><a href="${ontologyEditor.urls.objectPropertyHierarchy}" title="Object property hierarchy">Object property hierarchy</a></li>
+            <li role="listitem"><a href="${ontologyEditor.urls.propertyGroups}" title="Property groups">Property groups</a></li>
         </ul>
         
-        <h4><a href="${ontologyEditor.urls.ontologies}">Ontology list</a></h4>
+        <h4><a href="${ontologyEditor.urls.ontologies}" title="Ontology list">Ontology list</a></h4>
     </section>
 </#if>
\ No newline at end of file
diff --git a/webapp/web/templates/freemarker/body/siteAdmin/siteAdmin-siteConfiguration.ftl b/webapp/web/templates/freemarker/body/siteAdmin/siteAdmin-siteConfiguration.ftl
index ec170fa24..cb2ce4f2e 100644
--- a/webapp/web/templates/freemarker/body/siteAdmin/siteAdmin-siteConfiguration.ftl
+++ b/webapp/web/templates/freemarker/body/siteAdmin/siteAdmin-siteConfiguration.ftl
@@ -8,24 +8,24 @@
         
         <ul role="navigation">
             <#if siteConfig.internalClass?has_content>
-                <li role="listitem"><a href="${siteConfig.internalClass}">Institutional internal class</a></li>
+                <li role="listitem"><a href="${siteConfig.internalClass}" title="Institutional internal class">Institutional internal class</a></li>
             </#if>     
             
             <#if siteConfig.manageProxies?has_content>
-                <li role="listitem"><a href="${siteConfig.manageProxies}">Manage profile editing</a></li>
+                <li role="listitem"><a href="${siteConfig.manageProxies}" title="Manage profile editing">Manage profile editing</a></li>
             </#if>     
             
             <#if siteConfig.menuManagement?has_content>
-                <li role="listitem"><a href="${siteConfig.menuManagement}">Menu management</a></li>
+                <li role="listitem"><a href="${siteConfig.menuManagement}" title="Menu management">Menu management</a></li>
             </#if>      
             
             <#if siteConfig.siteInfo?has_content>
-                <li role="listitem"><a href="${siteConfig.siteInfo}">Site information</a></li>
+                <li role="listitem"><a href="${siteConfig.siteInfo}" title="Site information">Site information</a></li>
             </#if>
             
             <#if siteConfig.startupStatus?has_content>
                 <li role="listitem">
-                    <a href="${siteConfig.startupStatus}">Startup status</a>
+                    <a href="${siteConfig.startupStatus}" title="Startup status">Startup status</a>
                     <#if siteConfig.startupStatusAlert>
                         <img id="alertIcon" src="${urls.images}/iconAlert.png" width="20" height="20" alert="Error alert icon" />
                     </#if>
@@ -33,7 +33,7 @@
             </#if>   
             
              <#if siteConfig.userAccounts?has_content>
-                <li role="listitem"><a href="${siteConfig.userAccounts}">User accounts</a></li>
+                <li role="listitem"><a href="${siteConfig.userAccounts}" title="User accounts">User accounts</a></li>
              </#if>        
         </ul>
     </section>
diff --git a/webapp/web/templates/freemarker/body/termsOfUse.ftl b/webapp/web/templates/freemarker/body/termsOfUse.ftl
index f9859d97e..855397a53 100644
--- a/webapp/web/templates/freemarker/body/termsOfUse.ftl
+++ b/webapp/web/templates/freemarker/body/termsOfUse.ftl
@@ -9,10 +9,10 @@
     third parties, both individuals and organizations, commercial and otherwise. To the extent copyrightable,
     the information presented on the VIVO website and available as Resource Description Framework (RDF) data
     from VIVO at ${termsOfUse.siteHost} is intended for public use and is freely distributed under the terms of the
-    <a href="http://creativecommons.org/licenses/by/3.0/" target="_blank">Creative Commons CC-BY 3.0</a> license which allows you
+    <a href="http://creativecommons.org/licenses/by/3.0/" target="_blank" title="creative commons">Creative Commons CC-BY 3.0</a> license which allows you
     to copy, distribute, display and make derivatives of this information provided you give credit to 
     ${termsOfUse.siteHost}. Any non-copyrightable information is available to you under a
-    <a href="http://creativecommons.org/publicdomain/zero/1.0/"  target="_blank">CC0 waiver</a>.  However, source documents,
+    <a href="http://creativecommons.org/publicdomain/zero/1.0/"  target="_blank" title="cco waiver">CC0 waiver</a>.  However, source documents,
     images or web pages attached to or linked from VIVO may contain copyrighted information and should only be
     used or distributed under terms included with each source or in accordance with the principles of fair use.
     </p>
diff --git a/webapp/web/templates/freemarker/body/unrecognizedUser.ftl b/webapp/web/templates/freemarker/body/unrecognizedUser.ftl
index 105f5e199..8ff7a735e 100644
--- a/webapp/web/templates/freemarker/body/unrecognizedUser.ftl
+++ b/webapp/web/templates/freemarker/body/unrecognizedUser.ftl
@@ -11,5 +11,5 @@
     </p>
 
     <br/>
-    <a href="${urls.home}">Continue</a>
+    <a href="${urls.home}" title="continue">Continue</a>
 </section>
diff --git a/webapp/web/templates/freemarker/edit/forms/autoCompleteDatapropForm.ftl b/webapp/web/templates/freemarker/edit/forms/autoCompleteDatapropForm.ftl
index 62a77e00d..acf118276 100644
--- a/webapp/web/templates/freemarker/edit/forms/autoCompleteDatapropForm.ftl
+++ b/webapp/web/templates/freemarker/edit/forms/autoCompleteDatapropForm.ftl
@@ -63,7 +63,7 @@ $(document).ready(function() {
 
     <p class="submit">
         <input type="submit" id="submit" value="editConfiguration.submitLabel"/>
-        <span class="or"> or <a class="cancel" href="${editConfiguration.cancelUrl}">Cancel</a>
+        <span class="or"> or <a class="cancel" href="${editConfiguration.cancelUrl}" title="cancel">Cancel</a>
     </p>
 
     </form>
diff --git a/webapp/web/templates/freemarker/edit/forms/menuManagement-remove.ftl b/webapp/web/templates/freemarker/edit/forms/menuManagement-remove.ftl
index 4f965bb5a..05e46f78e 100644
--- a/webapp/web/templates/freemarker/edit/forms/menuManagement-remove.ftl
+++ b/webapp/web/templates/freemarker/edit/forms/menuManagement-remove.ftl
@@ -12,7 +12,7 @@
 
         <p>Are you sure you want to remove <em>${menuName}</em> menu item?</p>
 
-        <input type="submit" name="removeMenuItem" value="Remove menu item" class="submit" role="input" /> or <a class="cancel" href="${cancelUrl}">Cancel</a>
+        <input type="submit" name="removeMenuItem" value="Remove menu item" class="submit" role="input" /> or <a class="cancel" href="${cancelUrl}" title="cancel">Cancel</a>
     </form>
 </section>
 
diff --git a/webapp/web/templates/freemarker/edit/forms/menuManagement.ftl b/webapp/web/templates/freemarker/edit/forms/menuManagement.ftl
index ef7c8c363..e1aaec66f 100644
--- a/webapp/web/templates/freemarker/edit/forms/menuManagement.ftl
+++ b/webapp/web/templates/freemarker/edit/forms/menuManagement.ftl
@@ -46,7 +46,7 @@
             <p>Selected content type for the associated page</p>
             <p>
                 <span id="selectedContentTypeValue" name="selectedContentTypeValue">${associatedPage}</span>
-                <a href="#" id="changeContentType" name="changeContentType">Change content type</a>
+                <a href="#" id="changeContentType" name="changeContentType" title="change content type">Change content type</a>
             </p>
         </section>
         
@@ -92,7 +92,7 @@
             </ul>
         </section>
         
-        <input type="submit" name="submit-${menuAction}" value="Save changes" class="submit" role="input" /> or <a class="cancel" href="${cancelUrl}">Cancel</a>
+        <input type="submit" name="submit-${menuAction}" value="Save changes" class="submit" role="input" /> or <a class="cancel" href="${cancelUrl}" title="cancel">Cancel</a>
 
         <p class="requiredHint">* required fields</p>
     </form>
diff --git a/webapp/web/templates/freemarker/edit/forms/newIndividualForm.ftl b/webapp/web/templates/freemarker/edit/forms/newIndividualForm.ftl
index fc1b6746c..0606ae1fc 100644
--- a/webapp/web/templates/freemarker/edit/forms/newIndividualForm.ftl
+++ b/webapp/web/templates/freemarker/edit/forms/newIndividualForm.ftl
@@ -32,7 +32,7 @@
       <p class="submit">
           <input type="hidden" name = "editKey" value="${editKey}"/>
           <input type="submit" id="submit" value="Create ${typeName}"/>
-          <span class="or"> or <a class="cancel" href="${urls.base}/siteAdmin">Cancel</a>
+          <span class="or"> or <a class="cancel" href="${urls.base}/siteAdmin" title="cancel">Cancel</a>
       </p>
 
       <p id="requiredLegend" class="requiredHint">* required fields</p>
diff --git a/webapp/web/templates/freemarker/edit/forms/rdfsLabelForm.ftl b/webapp/web/templates/freemarker/edit/forms/rdfsLabelForm.ftl
index afd183a44..5779649ce 100644
--- a/webapp/web/templates/freemarker/edit/forms/rdfsLabelForm.ftl
+++ b/webapp/web/templates/freemarker/edit/forms/rdfsLabelForm.ftl
@@ -17,7 +17,7 @@
     
     <p class="submit">
         <input type="submit" id="submit" value="${submitLabel}" role="input"/>
-        or <a href="${cancelUrl}" class="cancel">Cancel</a>
+        or <a href="${cancelUrl}" class="cancel" title="cancel">Cancel</a>
     </p>
     
 </form>
diff --git a/webapp/web/templates/freemarker/lib/lib-properties.ftl b/webapp/web/templates/freemarker/lib/lib-properties.ftl
index b5a8ceaed..90ad6bea2 100644
--- a/webapp/web/templates/freemarker/lib/lib-properties.ftl
+++ b/webapp/web/templates/freemarker/lib/lib-properties.ftl
@@ -171,7 +171,7 @@ name will be used as the label. -->
     <#local verboseDisplay = property.verboseDisplay!>
     <#if verboseDisplay?has_content>       
         <section class="verbosePropertyListing">
-            <a class="propertyLink" href="${verboseDisplay.propertyEditUrl}">${verboseDisplay.localName}</a> 
+            <a class="propertyLink" href="${verboseDisplay.propertyEditUrl}" title="name">${verboseDisplay.localName}</a> 
             (<span>${property.type?lower_case}</span> property);
             order in group: <span>${verboseDisplay.displayRank};</span> 
             display level: <span>${verboseDisplay.displayLevel};</span>
@@ -200,7 +200,7 @@ name will be used as the label. -->
     <#-- Don't assume that if the mainImage property is populated, there is a thumbnail image (though that is the general case).
          If there's a mainImage statement but no thumbnail image, treat it as if there is no image. -->
     <#if (mainImage.statements)?has_content && thumbUrl?has_content>
-        <a href="${individual.imageUrl}"><img class="individual-photo" src="${thumbUrl}" title="click to view larger image" alt="${individual.name}" width="160" /></a>
+        <a href="${individual.imageUrl}" title="individual photo"><img class="individual-photo" src="${thumbUrl}" title="click to view larger image" alt="${individual.name}" width="160" /></a>
         <@editingLinks "${mainImage.localName}" mainImage.first() editable extraParams />
     <#else>
         <#local imageLabel><@addLinkWithLabel mainImage editable "Photo" extraParams /></#local>
diff --git a/webapp/web/templates/freemarker/page/partials/copyright.ftl b/webapp/web/templates/freemarker/page/partials/copyright.ftl
index 059479118..b0ca8758f 100644
--- a/webapp/web/templates/freemarker/page/partials/copyright.ftl
+++ b/webapp/web/templates/freemarker/page/partials/copyright.ftl
@@ -4,7 +4,7 @@
     <div class="copyright">
         &copy;${copyright.year?c}
         <#if copyright.url??>
-            <a href="${copyright.url}">${copyright.text}</a>
+            <a href="${copyright.url}" title="copyright">${copyright.text}</a>
         <#else>
             ${copyright.text}
         </#if>                 
diff --git a/webapp/web/templates/freemarker/page/partials/footer.ftl b/webapp/web/templates/freemarker/page/partials/footer.ftl
index efd30edec..2da9f78ba 100644
--- a/webapp/web/templates/freemarker/page/partials/footer.ftl
+++ b/webapp/web/templates/freemarker/page/partials/footer.ftl
@@ -18,7 +18,7 @@
 
     <#include "copyright.ftl">
 
-    All Rights Reserved. <a href="${urls.termsOfUse}">Terms of Use</a>
+    All Rights Reserved. <a href="${urls.termsOfUse}" title="terms of use">Terms of Use</a>
 
     <#include "version.ftl">
 </div>
diff --git a/webapp/web/templates/freemarker/page/partials/identity.ftl b/webapp/web/templates/freemarker/page/partials/identity.ftl
index fa20599f6..de755edf3 100644
--- a/webapp/web/templates/freemarker/page/partials/identity.ftl
+++ b/webapp/web/templates/freemarker/page/partials/identity.ftl
@@ -1,7 +1,7 @@
 <#-- $This file is distributed under the terms of the license in /doc/license.txt$ -->
 
 <header id="branding" role="banner">
-    <h1 class="vivo-logo"><a href="${urls.home}"><span class="displace">${siteName}</span></a></h1>
+    <h1 class="vivo-logo"><a href="${urls.home}" title="site name"><span class="displace">${siteName}</span></a></h1>
     <#if siteTagline?has_content>
         <em>${siteTagline}</em>
     </#if>
diff --git a/webapp/web/templates/freemarker/page/partials/menu.ftl b/webapp/web/templates/freemarker/page/partials/menu.ftl
index 356acce95..c5595ef95 100644
--- a/webapp/web/templates/freemarker/page/partials/menu.ftl
+++ b/webapp/web/templates/freemarker/page/partials/menu.ftl
@@ -3,7 +3,7 @@
 <nav role="navigation">
     <ul id="main-nav" role="list">
         <#list menu.items as item>
-            <li role="listitem"><a href="${item.url}" <#if item.active> class="selected" </#if>>${item.linkText}</a></li>
+            <li role="listitem"><a href="${item.url}" <#if item.active> class="selected" </#if> title="menu item">${item.linkText}</a></li>
         </#list>
     </ul>
 </nav>
diff --git a/webapp/web/templates/freemarker/page/partials/subMenuLinks.ftl b/webapp/web/templates/freemarker/page/partials/subMenuLinks.ftl
index 6191f4e60..639f802d0 100644
--- a/webapp/web/templates/freemarker/page/partials/subMenuLinks.ftl
+++ b/webapp/web/templates/freemarker/page/partials/subMenuLinks.ftl
@@ -6,5 +6,5 @@
 <#if urls.contact??>
     <li><a href="${urls.contact}">Contact Us</a></li>
 </#if> 
-<li><a href="http://www.vivoweb.org/support">Support</a></li>
-<li><a href="${urls.index}">Index</a></li>
\ No newline at end of file
+<li><a href="http://www.vivoweb.org/support" title="support">Support</a></li>
+<li><a href="${urls.index}" title="index">Index</a></li>
\ No newline at end of file
diff --git a/webapp/web/templates/freemarker/page/partials/version.ftl b/webapp/web/templates/freemarker/page/partials/version.ftl
index 4821efe65..4a515e64f 100644
--- a/webapp/web/templates/freemarker/page/partials/version.ftl
+++ b/webapp/web/templates/freemarker/page/partials/version.ftl
@@ -5,6 +5,6 @@
 <#-- Only show version info if user has access -->
 <#if user.hasRevisionInfoAccess>
     <div id="revision">
-        Version <a href="${version.moreInfoUrl}">${version.label}</a>
+        Version <a href="${version.moreInfoUrl}" title="version">${version.label}</a>
     </div>
 </#if>
\ No newline at end of file
diff --git a/webapp/web/templates/freemarker/widgets/widget-browse.ftl b/webapp/web/templates/freemarker/widgets/widget-browse.ftl
index 57e0fb0f1..e99388fe9 100644
--- a/webapp/web/templates/freemarker/widgets/widget-browse.ftl
+++ b/webapp/web/templates/freemarker/widgets/widget-browse.ftl
@@ -17,7 +17,7 @@
         <ul id="browse-classgroups" role="list">
         <#list vclassGroupList as group>
             <#if (group.individualCount > 0)>
-                <li role="listitem"><a href="${urls.currentPage}?classgroupUri=${group.uri?url}">${group.displayName?capitalize} <span class="count-classes">(${group.individualCount})</span></a></li>
+                <li role="listitem"><a href="${urls.currentPage}?classgroupUri=${group.uri?url}" title="group name">${group.displayName?capitalize} <span class="count-classes">(${group.individualCount})</span></a></li>
             </#if>
         </#list>
         </ul>
@@ -35,7 +35,7 @@
                  <ul id="classes-in-classgroup" role="list">
                      <#list classes as class>
                         <#if (class.individualCount > 0)>
-                            <li role="listitem"><a href="${urls.currentPage}?classgroupUri=${classGroup.uri?url}&vclassUri=${class.uri?url}">${class.name} <span class="count-individuals"> (${class.individualCount})</span></a></li>
+                            <li role="listitem"><a href="${urls.currentPage}?classgroupUri=${classGroup.uri?url}&vclassUri=${class.uri?url}" title="class name">${class.name} <span class="count-individuals"> (${class.individualCount})</span></a></li>
                         </#if>
                      </#list>
                  </ul>
@@ -54,7 +54,7 @@
          
         <ul>
             <#list individualsInClass as ind>
-                <li><a href="${urls.base}/individual?uri=${ind.uri?url}">${ind.name}</a></li>
+                <li><a href="${urls.base}/individual?uri=${ind.uri?url}" title="individual name">${ind.name}</a></li>
             </#list>
         </section>
 </#macro>
diff --git a/webapp/web/templates/freemarker/widgets/widget-login.ftl b/webapp/web/templates/freemarker/widgets/widget-login.ftl
index 34dd80d25..a9d3fcbc7 100644
--- a/webapp/web/templates/freemarker/widgets/widget-login.ftl
+++ b/webapp/web/templates/freemarker/widgets/widget-login.ftl
@@ -20,7 +20,7 @@
         <noscript>
             <section id="error-alert">
                 <img src="${urls.images}/iconAlertBig.png" alt="Alert Icon"/>
-                <p>In order to edit content, you'll need to enable JavaScript. Here are the <a href="http://www.enable-javascript.com">instructions for enabling JavaScript in your web browser</a>.</p>
+                <p>In order to edit content, you'll need to enable JavaScript. Here are the <a href="http://www.enable-javascript.com" title="java script instructions">instructions for enabling JavaScript in your web browser</a>.</p>
             </section>
         </noscript>
     </#if>
@@ -40,7 +40,7 @@
        
         <form role="form" id="login-form" action="${formAction}" method="post" name="login-form" />
             <#if externalAuthUrl??>
-                <p class="external-auth"><a class="blue button" href="${externalAuthUrl}">${externalAuthName}</a></p>
+                <p class="external-auth"><a class="blue button" href="${externalAuthUrl}" title="external authentication name">${externalAuthName}</a></p>
                 <p class="or-auth">or</p>
 
                 <h3 class="internal-auth" >Log in using your ${siteName} account</h3>
@@ -57,8 +57,8 @@
             <input class="checkbox-remember-me" name="remember-me" type="checkbox" value="" />  
             <label class="label-remember-me" for="remember-me">Remember me</label> -->
             <#-- mb863: forgot password and request an account won't be part of VIVO r1.2
-            <p class="forgot-password"><a href="#">Forgot your password?</a></p>
-            <p class="request-account"><a class="blue button" href="#">Request an account</a> </p> -->
+            <p class="forgot-password"><a href="#" title="forgot password">Forgot your password?</a></p>
+            <p class="request-account"><a class="blue button" href="#" title="request an account">Request an account</a> </p> -->
         </form>
 
     </section><!-- #log-in -->
@@ -83,7 +83,7 @@
                 <label for="confirmPassword">Confirm Password</label>
                 <input id="confirmPassword" name="confirmPassword" class="text-field" type="password" required />
                 
-                 <p class="submit-password"><input name="passwordChangeForm" class="green button" type="submit" value="Log in"/> <span class="or">or <a class="cancel" href="${cancelUrl}">Cancel</a></span></p>
+                 <p class="submit-password"><input name="passwordChangeForm" class="green button" type="submit" value="Log in"/> <span class="or">or <a class="cancel" href="${cancelUrl}" title="cancel">Cancel</a></span></p>
             </form>
     </section>
 </#macro>